Organization Nuffield Council on Bioethics

University of Edinburgh

Royal Edinburgh Hospital

Institute of Molecular Medicine

National Academy of Sciences

British Association for the Advancement of Science (President, 1992-1993)

Royal Society (Fellow)

Academy of Medical Sciences (Fellow)

King's College London

Queen Mary College, University of London

British Association for the Advancement of Science (president)

Medical Research Council, London

Wellcome Trust/Cancer Research UK Gurdon Institute

Warnock Committee